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Santalum haleakalae var. lanaiense

(Rock) Harbaugh (2010)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Santalopsida Order: Santalales Family: Santalaceae Genus: Santalum

‘iliahi [iliahi], lā‘au ala, sandalwood

Description

Key Characters:

Growth Form: Small trees, evergreen, single or multiple stems; 1–6 m tall; apparently hemiparasitic on roots of other plants.

Stems: Older trees readily spread vegetatively via root sprouts when surface roots are exposed or damaged, sometimes forming small stands.

Roots:

Leaves: Leaves simple. Opposite. Blades narrowly to broadly ovate, to elliptic (rarely conduplicate–falcate), (3.4–)4.4–6.8(–8.4) cm long × (1.0–)1.6–3.1(–4.9) cm wide. Apex acute or obtuse (rarely acuminate). Base cuneate. More glaucous Lower surfacely, surfaces paler Lower surfacely, Light, evergreen, olive, to dark green, chartaceous to coriaceous. Margins flat to slightly recurved. Leaf lateral veins apparent on both surfaces of the leaf, yellowish to dark purple. Petioles (3.9–) 7.4–15.4 (–21.7) mm long. Stipules absent.

Flowers: Flowers in inflorescences fairly dense to dense, terminal or axillary. Flowers bisexual (perfect), weakly fragrant, reddish to cream-colored in bud, longer than wide. Pedicels 2–4 mm long, bracts 1–4 mm in length, persistent until flowering. Calyx (sepals) absent. Corolla of 4 (rarely 5) petals; petals deep pink to red throughout, or with white to pink interiors; concave with thickened margins, papillate, (2.0–)3.0–4.4(–4.8) mm long × (1.8–)1.9–2.7(–3.8) mm wide; floral tubes (3.4–)4.2–8.0(–11.2) mm long × (1.9–)2. Stamens 4, hairs proximal to the filaments (rarely hairs distal). Ovary partly inferior (approx. ⅔); drupe elliptic, with subapical ring; stigma lobes 3.

Fruit: Drupes black or purplish black at maturity; 10–16 mm long; with a subapical receptacular ring. Seeds muricate; with pointed beak on one end.

Ploidy:

Habitat: Exposed or semishaded scrublands.

Elevation Range: 800–1,300 m.

Historical Distribution

Uses and Culture

USES

Natural History

Statewide Status

Endemic

Island Status

Molokai Endemic
Lana'i Endemic
Maui Endemic

Dispersal Agents


Pollinators

Bibliography

Name Published In: Syst. Bot. 35: 834 (2010)

Other References

Wagner et al. 1990:1221 (L, M [as S. freycinetianum var. lanaiense]); Harbaugh et al. 2010:834 (COMBNOV, Syn. S. freycinetianum var. lanaiense = S. haleakalae var. lanaiense, add Mo, KEY, DESCR)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Forbes, C.N. 327 Lanai BISH 1917-09-01
2 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Munro, G.C. 951 Lanai BISH 1929-01-01
3 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Hughes, G.D. 56 Molokai BISH 1992-09-09
4 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Pratt, T.K. Molokai BISH 1972-07-02
5 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Munro, G.C. 950 Lanai BISH 1929-01-01
6 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Stemmermann, R.L. 812 Lanai BISH 1976-04-30
7 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details in company with Psychotria, Zanthoxylum, Dracaena, Myrsine, Osmanthus, Diospyros, Metrosideros. Preserved_Specimen Kepler, A.K. Molokai BISH 1980-08-06
8 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details In Metrosideros-Nestegis lowland mesic forest with Dicranopteris, Wikstroemia, Broussaisia, Cibotium, Myrsine, Coprosma, Hedyotis, Cheirodendron, Clermontia, Cyanea, Labordia, Psychotria, Pittosporum, Myrica faya, Pisonia, Styphelia. Preserved_Specimen Perlman, S. 6375 Lanai BISH 1987-08-26
9 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Very open 'ohia forest w/ 'a'ali'i & ground cover of uluhe & wawae'iole. Preserved_Specimen Davis, J. 790 Molokai BISH 1982-05-22
10 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Lowland Ohia Mixed Mesic Forest at 3550 ft in a gulch between two southern ridges with Metrosideros, Coprosma, Dodonaea, Myrsine, Wikstroemia, Charpentiera, Alyxia, Hedyotis, Vaccinium, Cyrtandra, Astelia, Dubautia, Pelea, Smilax, Pouteria, Nestegis and native ferns and sedges Preserved_Specimen Welton, P. 1507 Maui BISH 1992-01-11
11 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Thicket on moist ridge. Preserved_Specimen St.John, H. 18712 Lanai BISH 1938-04-09
12 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Near ridge top on steep, rocky N facing slope. Vegetation: Dodonaea, Leptecophylla, Metrosideros, Myrsine, Coprosma, Wikstroemia, Dubautia, Alyxia, Bidens, Osteomeles, Carex, Eragrostis, Pteridium. Preserved_Specimen Oppenheimer, H.L. 120609 Maui BISH 2006-12-06
13 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Open dry windswept forest of Osmanthus. Preserved_Specimen St.John, H. 22625 Lanai BISH 1947-04-05
14 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Dry forest. Preserved_Specimen Little Jr., E.L. 31251 Lanai BISH 1976-09-17
15 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Metrosideros, Diospyros, Pleomele, Styphelia, Coprosma foliosa, Nestegis, Melicope volcanica, Dodonaea, Lysimachia remyi, Doodia kunthiana, Wikstroemia, Pittosporum, Lantana, Buddleja, Psychotria Preserved_Specimen Perlman, S. 18985 Maui BISH 2004-05-26
16 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Open forest of Osmanthus, Dodonaea, Metrosideros etc., with grassy undergrowth. Preserved_Specimen Stauffer, H.U. 5892 Maui BISH 1964-04-18
17 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Metrosideros polymorpha var. glaberrima, Dicranopteris linearis, Dodonaea viscosa, Machaerina sp., Dianella sandwicensis, Hedyotis terminalis and Palhinhaea cernua Preserved_Specimen Oppenheimer, H.L. 20011 Maui BISH 2000-02-23
18 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Solitary tree in open area past the edge of the forest, on grassy ridge crest. Preserved_Specimen Lau, J. 2868 Lanai BISH 1986-10-05
19 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Metrosideros forest. Preserved_Specimen Little Jr., E.L. 31233 Lanai BISH 1976-09-17
20 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Both sides of ridge, on upper slopes. Vegetation: Metrosideros, Dicranopteris, Psychotria, Vaccinium, Pteridium, Leptecophylla, Machaerina, Ilex, Coprosma, Kadua, Perrottetia, Andropogon, Axonopus. Preserved_Specimen Oppenheimer, H.L. 80902 Molokai BISH 2009-08-03
21 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Small tree in windswept elfin forest. With Gardenia brighami, Osmanthus, Planchonella, etc. In dry stony soil. Preserved_Specimen Stone, B.C. 883 Lanai BISH 1955-07-03
22 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Mesic ohia forest with other native trees and a native shrub, matted-fern understory. On exposed ridge in full sun. Preserved_Specimen Warshauer, F.R. 2332 Molokai BISH 1979-06-21
23 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Low wet forest with Metrosideros dominant on lower slopes and valley bottom, with Styphelia/Dodonaea shrubland on upper slopes and ridgetops. Preserved_Specimen Lorence, D.H. 5612 Molokai BISH 1987-10-21
24 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Remnant of dry forest with more than a score of native trees on less than 50 acres. Preserved_Specimen Sohmer, S.H. 6603 Maui BISH 1970-08-20
25 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details On steep S facing slopes; few trees scattered in this area mid-to lower slope. Vegetation: Metrosideros, Cheirodendron, Coprosma, Broussaisia, Zanthoxylum, Clermontia, Alyxia, Freycinetia, Microlepia, Perrottetia, Diplazium, Ilex, Sadleria. Preserved_Specimen Oppenheimer, H.L. 50710 Maui BISH 2007-05-07
26 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details With Styphelia, Acacia & koa. Preserved_Specimen Medeiros, A.C. 540 Maui BISH 1983-09-27
27 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Remnant dryland sclerophyll forest. Preserved_Specimen Wagner, W.L. 5463 Maui BISH 1984-10-07
28 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Remnant of dry forest with more than a score of native trees on less than 50 acres. Preserved_Specimen Sohmer, S.H. 6603 Maui BISH 1970-08-20
29 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Dry-land native forest. Preserved_Specimen Desha, A. Lanai BISH 1963-05-04
30 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details In open Metrosideros/Dicranopteris Forest. Vegetation: Cheirodendron, Ilex, Broussaisia, Myrsine, Dodonaea, Scaevola, Psychotria, Pteridium, Sadleria. Preserved_Specimen Oppenheimer, H.L. 120612 Maui BISH 2006-12-06
31 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Open forest of Osmanthus, Dodonaea, Metrosideros etc., with grassy undergrowth. Preserved_Specimen Stauffer, H.U. 5892 Maui BISH 1964-04-18
32 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Mesic ohia forest with other native trees and native shrub-exotic grass and shrub understory. Preserved_Specimen Warshauer, F.R. 3003 Molokai BISH 1980-01-01
33 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Sloping patch of dryish scrub forest. Preserved_Specimen Degener, O. 35806 Lanai BISH 1964-02-01
34 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Open shrubby rain-forest. Preserved_Specimen Degener, O. 24255 Lanai BISH 1957-08-15
35 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Open shrubby rain-forest at top. Preserved_Specimen Degener, O. 24257 Lanai BISH 1957-08-13
36 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details On steep gulch slope. Preserved_Specimen Hobdy, R.W. 1722 Maui BISH 1982-12-15
37 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Montane open Metrosideros with native shrubs and Dicranopteris. Preserved_Specimen Pratt, L.W. 1145 Molokai BISH 1982-05-22
38 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details On very steep slope. Associated vegetation: Acacia, Metrosideros, Tetraplasandra, Melicope, Ilex, Myrsine, Cheirodendron, Wikstroemia, Dodonaea, Alyxia, Smilax, Freycinetia, Doodia, Sadleria, Microlepia. Preserved_Specimen Oppenheimer, H.L. 90604 Maui BISH 2006-09-07
39 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Vegetation: Metrosideros, Dodonaea, Dicranopteris, Diplopterygium, Coprosma, Sadleria, Ilex, Myrsine, Vaccinium, Miconia crenata, Tibouchina, Rubus, Peperomia, Psilotum. Preserved_Specimen Oppenheimer, H.L. 22340 Maui BISH 2023-02-14
40 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Locally common in dry open sclerophyllous forest on aa lava flow. Preserved_Specimen Fosberg, F.R. 41515 Maui BISH 1961-03-14
41 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Dry forest on lava. Preserved_Specimen Little Jr., E.L. 31131 Maui BISH 1976-08-26
42 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Small tree in remnant of native dry-land forest. Preserved_Specimen Herbst, D.R. 5789 Lanai BISH 1976-03-31
43 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Mesic native tree community with scattered exotic trees and exotic grass-shrub understory. Preserved_Specimen Jacobi, J.D. 1262 Lanai BISH 1979-04-05
44 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details On dry a`a lava in open, native forest. Preserved_Specimen Herbst, D.R. 728 Maui BISH 1967-11-12
45 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Closed to open Metrosideros mesic forest and riparian vegetation. W/ matting ferns of Dicranopteris and Diplopterygium with Freycinetia, Labordia, Xylosma, Antidesma, Santalum, Pleomele, Pouteria, Melicope volcanica, Kadua acuminata, K. affinis, Tetraplasandra oahuensis. Preserved_Specimen Wood, K.R. 12750 Lanai BISH 2008-01-09
46 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Mesic native shrub community with scattered ohia trees Preserved_Specimen Stemmermann, R.L. 3896 Molokai BISH 1979-01-01
47 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Open dry windswept forest of Osmanthus. Preserved_Specimen St.John, H. 22625 Lanai BISH 1947-04-05
48 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Stemmermann, R.L. 819 Lanai BISH 1976-04-30
49 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Stemmermann, R.L. 755 Maui BISH 1976-06-26
50 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Rock, J.F.C. 10061 Lanai BISH 1910-07-01
51 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Stemmermann, R.L. 811 Lanai BISH 1976-04-30
52 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Munro, G.C. 679 Lanai BISH 1919-02-18
53 Santalum haleakalae var. lanaiense West Maui, Kapunakea. Preserved_Specimen Hughes, G.D. 611 Maui BISH 1993-05-11
54 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Munro, G.C. 43 Lanai BISH 1913-11-04
55 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Munro, G.C. 200 Lanai BISH 1913-11-04
56 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Hobdy, R.W. 2620 Molokai BISH 1986-08-19
57 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Munro, G.C. 21 Lanai BISH 1927-05-28
58 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Stemmermann, R.L. 2149 Maui BISH 1977-06-01
59 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Rock, J.F.C. 27002 Maui BISH 1953-02-01
60 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Stemmermann, R.L. 2149 Maui BISH 1977-06-01
61 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Munro, G.C. 949 Lanai BISH 1929-01-01
62 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Medeiros, A.C. 196 Maui BISH 1981-12-18
63 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Munro, G.C. 946 Lanai BISH 1929-08-21
64 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Rock, J.F.C. 27002 Maui BISH 1953-02-01
65 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Stemmermann, R.L. 813 Lanai BISH 1976-04-30
66 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Wagner, W.L. 4933 Molokai BISH 1983-08-01
67 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Rock, J.F.C. 8659 Maui BISH 1910-11-01
68 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Hobdy, R.W. 1163 Molokai BISH 1981-10-26
69 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Munro, G.C. 211 Lanai BISH 1927-09-09
70 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Munro, G.C. 497 Lanai BISH 1921-06-01
71 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Hobdy, R.W. 2619 Molokai BISH 1986-08-19
72 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Forbes, C.N. 114 Lanai BISH 1913-06-01
73 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Stemmermann, R.L. 2149 Maui BISH 1977-06-01
74 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Forbes, C.N. 229 Lanai BISH 1913-06-01
75 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Russell, C.A. Molokai BISH 1986-10-16
76 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Russell, C.A. Molokai BISH 1987-10-22
77 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Montgomery, S.L. Lanai BISH 1971-06-11
78 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Russell, C.A. Molokai BISH 1987-10-22
79 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Russell, C.A. Molokai BISH 1987-10-22
80 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Munro, G.C. Lanai BISH 1916-07-20
81 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Rock, J.F.C. Lanai BISH 1912-10-25
82 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Munro, G.C. Lanai BISH 1920-06-28
83 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Medeiros, A.C. 182 Maui BISH 1981-10-01
84 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Hobdy, R.W. 3303 Maui BISH 1991-06-04
85 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen St.John, H. 26863 Maui BISH 1972-06-30
86 Santalum haleakalae var. lanaiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Munro, G.C. 678 Lanai BISH 1919-04-02
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